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Invertebrate Identification

Queensland, Australia.

Assisted research project, looking for undescribed species in the gastropod families: Stenothyridae, Iravadiidae, Calopiidae.


American Malacological Society

Malacological Society of Australasia

Conchological Society of Great Britain and Ireland

The Ecological Consultants Association of New South Wales

Malacological Society of London

National Shellfisheries Association

Royal Zoological Society of New South Wales

Member of the IUCN SSC Mollusc Specialist Group

Research Associate:

Field Museum of Natural History, Chicago, IL.

 Florida Museum of Natural History, Gainesville, FL.

Anodonta californiensis,

photo: Katharine Hollister, Invertebrate Identification


For inclusion in a Field Guide to the Freshwater and Terrestrial Molluscs of the Sacramento-Pit River area, for the California Fish & Game Service.

* Current paper awaiting acceptance.

Clark, S.A. 2019. Amnicolidae Tryon, 1862. In: Freshwater mollusks of the world. A distribution atlas. Eds. Lydeard, C. and Cummings, K.S. Johns Hopkins University Press, Baltimore, Maryland, USA, pp. 90-93.


Clark, S.A. 2019. Cochliopidae Tryon, 1866. In: Freshwater mollusks of the world. A distribution atlas. Eds. Lydeard, C. and Cummings, K.S. Johns Hopkins University Press, Baltimore, Maryland, USA, pp. 104-108.


Clark, S.A. 2019. Lithoglyphidae Tryon, 1866. In: Freshwater mollusks of the world. A distribution atlas. Eds. Lydeard, C. and Cummings, K.S. Johns Hopkins University Press, Baltimore, Maryland, USA, pp. 118-121.


Clark, S.A. 2019. Stenothyridae Tryon, 1866. In: Freshwater mollusks of the world. A distribution atlas. Eds. Lydeard, C. and Cummings, K.S. Johns Hopkins University Press, Baltimore, Maryland, USA, pp. 131-133.


Keenan, S.W., Audrey T. Paterson, A.T., Niemiller, M.L., Slay, M.E., Clark, S.A. and Engel, A.S. 2017. Observations of the first stygobiont snail (Hydrobiidae, Fontigens sp.) in Tennessee. Proceedings of the 17th International Congress of Speleology 2017:91-94.


Campbell, D.C., Clark, S.A. and Lydeard, C. 2017. Phylogenetic analysis of the Lancinae (Gastropoda, Lymnaeidae) with a description of the U.S. federally endangered Banbury Springs lanx. ZooKeys 663:107-132.


Ponder, W.F., Hallan, A., Shea, M. and Clark, S.A. 2016. Australian Freshwater Molluscs. The snails and bivalves of Australian inland waters. Interactive key


Johannes, E.J. and Clark, S.A. 2016. Freshwater mollusc declines, local extinctions and introductions in five northern California streams. Tentacle 24:22-25.


Campbell, D., Clark, S.A., Johannes, E., Lydeard, C. and Frest, T. 2016. Molecular phylogenetics of the freshwater gastropod genus Juga (Cerithioidea: Semisulcospiridae). Biochemical Systematics and Ecology 65:158-170.


Gerber, J. and Clark, S.A. 2015. First record of the predatory land snail Streptostele (Tomostele) musaecola (Pulmonata: Streptaxidae) in the continental United States. American Conchologist 43(4):26-28.


Hauk, A., Clark, S.A., McCravy, K.W., Jenkins, S.E. and Lydeard, C. 2015. A Survey of Terrestrial Gastropods of the Alice L. Kibbe Life Science Station in West-Central Illinois. Northeastern Naturalist 22(2):299-306.


2014. Bieler, R., Mikkelsen, P.M., Timothy M. Collins, T.M., Glover, E.A., González, V.L., Daniel L. Graf, D.L., Harper, E.M., John Healy, J., Kawauchi, G.Y., Sharma, P.P., Staubach, S., Strong, E.E., Taylor, J.D., Tëmkin, I., Zardus, J.D., Clark, S., Guzmán, A., McIntyre, E., Sharp, P. and Giribet, G.  Investigating the Bivalve Tree of Life – an exemplar-based approach combining molecular and novel morphological characters. Invertebrate Systematics 28(1):32-115.

Clark, S.A. 2009. Revision of the genus Posticobia (Mollusca: Caenogastropoda: Rissooidea: Hydrobiidae s.l.) from Australia and Norfolk Island. Malacologia 51(2):319-341.

Clark, S.A. 2009. A review of the land snail genus Meridolum (Gastropoda: Camaenidae) from central New South Wales, Australia. Molluscan Research 29(2):61-120.


Ó Foighil, D., Lee, T., Campbell, D.C. and Clark, S.A. 2009. All voucher specimens are not created equal: a cautionary tale involving North American pleurocerid gastropods. Journal of Molluscan Studies 75(3):305-306.


Waggoner, J., Clark, S.A., Perez, K.E. and Lydeard, C. 2006. A survey of terrestrial gastropods of the Sipsey Wilderness (Bankhead National Forest), Alabama. Southeastern Naturalist 5(1):57-68.


Ponder, W.F., Clark, S.A., Eberhard, S. and Studdert, J.B. 2005. A radiation of hydrobiid snails in the caves and streams at Precipitous Bluff, southwest Tasmania, Australia (Mollusca: Caenogastropoda: Rissooidea: Hydrobiidae s.l.). Zootaxa 1074:1-66.


Perez, K.E., Ponder, W.F., Colgan, D.J., Clark, S.A. and Lydeard, C. 2005. Molecular phylogeny and biogeography of spring-associated hydrobiid snails of the Great Artesian Basin, Australia. Molecular Phylogenetics and Evolution 34(3):545-556.


Clark, S.A. 2005. Systematics, spatial analysis and conservation genetics of Meridolum corneovirens and related forms (Gastropoda: Camaenidae) from the Sydney Region of Australia. Ph.D. Thesis, University of Western Sydney, Richmond, Sydney, New South Wales. 256 pp.


Lydeard, C., Cowie, R.H., Ponder, W.F., Bogan, A.E., Bouchet, P., Clark, S.A., Cummings, K.S., Frest, T.J., Gargominy, O., Herbert, D.G., Hershler, R., Perez, K.E., Roth, B., Seddon, M., Strong, E.E., Thompson, F.G. 2004. The global decline of nonmarine mollusks. Bioscience 54(4):321-330.


Clark, S.A. 2004. Native snails in an urban environment – conservation from the ground up. In Urban wildlife: more than meets the eye, eds Lunney, D. and Burgin, S., Royal Zoological Society of New South Wales, Mosman, NSW, Australia, 78-81.


Clark, S.A.; Miller, A.C. and Ponder, W.F. 2003. Revision of Austropyrgus (Gastropoda: Hydrobiidae); a morphostatic radiation of freshwater gastropods in south-eastern Australia. Records of the Australian Museum, Supplement 28:1-109.


Clark, S.A. and Richardson, B.J. 2002. Spatial analysis of genetic variation as a rapid assessment tool in the conservation management of narrow range endemics. Invertebrate Systematics 16(4):583-587.


Ponder, W.F., Clark, S.A. and Dallwitz, M.J. 2000. Freshwater and Estuarine Molluscs. An interactive, illustrated key for New South Wales. CD-ROM, CSIRO Publishing, Melbourne, Australia.


Miller, A.C., Ponder, W.F. and Clark, S.A. 1999. Freshwater snails of the genera Fluvidona and Austropyrgus (Gastropoda, Hydrobiidae) from northern New South Wales and southern Queensland, Australia. Invertebrate Taxonomy 13(3):461-493.


Ponder, W.F., Clark, S.A. and Miller, A.C. 1999. A new genus and two new species of Hydrobiidae (Mollusca: Gastropoda: Caenogastropoda) from south Western Australia. Proceedings of the Royal Society of Western Australia 82(3):109-120.


Clark, S.A. 1997. Taxonomy and biology of Posticobia (Gastropoda: Hydrobiidae). M.Sc. Thesis, Macquarie University, North Ryde, Sydney, New South Wales. 199 pp.


Ponder, W.F., Colgan, D.J., Terzis, T., Clark, S.A. and Miller, A.C. 1996. Three new morphologically and genetically determined species of hydrobiid gastropods from Dalhousie Springs, northern South Australia, with the description of a new genus. Molluscan Research 17:49-106.


Journal and Peer Reviewed Publications:

Stephanie Clark, Ph.D (S.A. Clark)
Malacologist, Taxonomist, Conchologist














United States, Canada, Australasia, Oceania.

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